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<title>August 4 Friday Evening Lecture to Discuss Stem Cells in Thorn Development and Tackling the Challenges of Citrus Agriculture</title>
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<p class="MsoNormal" style="line-height:150%"><span style="font-size:8.5pt;line-height:150%;font-family:"Helvetica Neue";color:#656565">Irish is the Eaton Professor of Molecular, Cellular and Developmental Biology, and Professor of Ecology and Evolutionary
Biology, at Yale University.<o:p></o:p></span></p>
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<p class="MsoNormal" style="line-height:150%"><strong><span style="font-size:12.0pt;line-height:150%;font-family:Helvetica;color:#F2F2F2">FRIDAY, AUGUST 4</span></strong><b><span style="font-size:12.0pt;line-height:150%;font-family:Helvetica;color:#F2F2F2"><br>
<strong><span style="font-family:Helvetica">8:00 PM | Cornelia Clapp Auditorium</span></strong></span></b><span style="font-size:12.0pt;line-height:150%;font-family:Helvetica;color:#F2F2F2"><br>
<em><span style="font-family:Helvetica">Porter Lecture</span></em></span><span style="font-size:10.5pt;line-height:150%;font-family:Helvetica;color:#F2F2F2"><br>
<br>
</span><span class="tweddescription"><b><span style="font-size:12.0pt;line-height:150%;font-family:Helvetica;color:#F2F2F2">"</span></b></span><span class="tweddescription"><b><span style="font-size:10.5pt;line-height:150%;font-family:Helvetica;color:#F2F2F2">Thorny
Questions: Citrus Development and Disease</span></b></span><span class="tweddescription"><b><span style="font-size:12.0pt;line-height:150%;font-family:Helvetica;color:#F2F2F2">"</span></b></span><span style="font-size:10.5pt;line-height:150%;font-family:Helvetica;color:#F2F2F2"><br>
<br>
<strong><span style="font-family:Helvetica">Vivian Irish, </span></strong>Yale University<br>
<br>
Offered in-person and virtually<br>
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<p class="MsoNormal" style="line-height:150%"><span style="font-size:10.5pt;line-height:150%;font-family:Helvetica;color:#F2F2F2">Because plants are sessile and can’t escape from predators, many plant species have evolved exquisite defensive systems, ranging
from chemical defenses to physical deterrents. Thorns, prickles, and spines all act as weapons against herbivores, but each arises from a distinct developmental origin. Thorns represent a fascinating variation on how stem cells are regulated. Irish's work
is focused on the gene network necessary for thorn development in citrus species to help tackle some of the challenges facing citrus agriculture today. <a href="https://nam02.safelinks.protection.outlook.com/?url=https%3A%2F%2Fmbl.us14.list-manage.com%2Ftrack%2Fclick%3Fu%3D9cefa84d03e46926e3394b4d2%26id%3Db0f22de0df%26e%3Dcc1a53fb05&data=05%7C01%7Csemco%40whoi.edu%7Cd8be5edec9234f67372008db943db7b8%7Cd44c5cc6d18c46cc8abd4fdf5b6e5944%7C0%7C0%7C638266765018604638%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=459w7qG1Qbsk30Rc7C7Pgp0WNsrG2Ks5BtOVfIwjaXI%3D&reserved=0" originalSrc="https://mbl.us14.list-manage.com/track/click?u=9cefa84d03e46926e3394b4d2&id=b0f22de0df&e=cc1a53fb05" shash="gp1SJjhH2oDbctKlpP2OVCTY9d4teyPuDqRZYoMFs4qo6xpaGxwI+zUbKEHV6CAZb0b4Sf6W3ogHB5DwpJ6uuCkD5iMk8XqBRFpznsepROVGBnXZPxm3HJqEaMA7iNiY+9iSXIPy+2n7uaT8iAuv+bwLBHZwDzKLIdXRiZBwvdU=" target="_blank"><b><span style="color:white">Read
more >></span></b></a><o:p></o:p></span></p>
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<td width="600" valign="top" style="width:6.25in;padding:0in 0in 0in 0in;max-width:100%;min-width: 100%;-ms-text-size-adjust: 100%;-webkit-text-size-adjust: 100%">
<table class="MsoNormalTable" border="0" cellspacing="0" cellpadding="0" align="left" width="100%" style="width:100.0%;border-collapse:collapse;-ms-text-size-adjust: 100%;-webkit-text-size-adjust: 100%;word-break:break-word">
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<td valign="top" style="padding:0in 13.5pt 6.75pt 13.5pt;-ms-text-size-adjust: 100%;-webkit-text-size-adjust: 100%">
<p class="MsoNormal" align="center" style="text-align:center;line-height:150%"><span style="font-size:10.5pt;line-height:150%;font-family:"Helvetica Neue";color:black"><a href="https://nam02.safelinks.protection.outlook.com/?url=https%3A%2F%2Fmbl.us14.list-manage.com%2Ftrack%2Fclick%3Fu%3D9cefa84d03e46926e3394b4d2%26id%3Dee49ec774c%26e%3Dcc1a53fb05&data=05%7C01%7Csemco%40whoi.edu%7Cd8be5edec9234f67372008db943db7b8%7Cd44c5cc6d18c46cc8abd4fdf5b6e5944%7C0%7C0%7C638266765018760850%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=J4leAiEfMDMyYsB%2FPOx70TQgAHqD1JlIIroaCxuNVsk%3D&reserved=0" originalSrc="https://mbl.us14.list-manage.com/track/click?u=9cefa84d03e46926e3394b4d2&id=ee49ec774c&e=cc1a53fb05" shash="R9fyoANBXyHvTKRCjuNzDOslyT+MzXtP30GUrSPzbfMThoP5eHxpwWk2ChpPtMQpffPmtJr20W3lRCF4a6y9boGfg19yuR9i+bnWWrJGyyEYt9IC2i9PuYEOgmL9u8tKKbhMUVkPkTDAa8iGyP8Os5POs0WB8ebtcvP17AG0vHU=" target="_blank"><strong><span style="font-size:12.0pt;line-height:150%;font-family:"Helvetica Neue";color:#007396">or
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<table class="MsoNormalTable" border="0" cellspacing="0" cellpadding="0" align="left" style="border-collapse:collapse;-ms-text-size-adjust: 100%;-webkit-text-size-adjust: 100%">
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<p class="MsoNormal" align="center" style="text-align:center"><a href="https://nam02.safelinks.protection.outlook.com/?url=https%3A%2F%2Fmbl.us14.list-manage.com%2Ftrack%2Fclick%3Fu%3D9cefa84d03e46926e3394b4d2%26id%3D3e43ab55fd%26e%3Dcc1a53fb05&data=05%7C01%7Csemco%40whoi.edu%7Cd8be5edec9234f67372008db943db7b8%7Cd44c5cc6d18c46cc8abd4fdf5b6e5944%7C0%7C0%7C638266765018760850%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=%2B50MapvpuKmtbBDP4GGNFSthYbHFbIZumArg%2BleZgmw%3D&reserved=0" originalSrc="https://mbl.us14.list-manage.com/track/click?u=9cefa84d03e46926e3394b4d2&id=3e43ab55fd&e=cc1a53fb05" shash="DxG6OSfIwHprnSzB22m7Y7p+zNZ6+HwJQSBCoTOh/s05iSvcNtZHj/GQZgymzZscUYGB/tBZZeRh8xP+MgI74Xmrz2D8IPM05u5PUXsbLleALwt7n4gqdHWDvFFJYX80hmiGCXVpuX3UvyiaXdgNoLAPeuII4tM68Xhovtuldks=" target="_blank"><span style="color:windowtext;text-decoration:none"><span style="color:blue"><img border="0" width="24" height="24" style="width:.25in;height:.25in" id="_x0000_i1029" src="https://cdn-images.mailchimp.com/icons/social-block-v2/outline-light-twitter-48.png" alt="Twitter"></span></span></a><o:p></o:p></p>
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<td style="padding:3.75pt 7.5pt 3.75pt 6.75pt;-ms-text-size-adjust: 100%;-webkit-text-size-adjust: 100%">
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<td width="24" style="width:.25in;padding:0in 0in 0in 0in;-ms-text-size-adjust: 100%;-webkit-text-size-adjust: 100%">
<p class="MsoNormal" align="center" style="text-align:center"><a href="https://nam02.safelinks.protection.outlook.com/?url=https%3A%2F%2Fmbl.us14.list-manage.com%2Ftrack%2Fclick%3Fu%3D9cefa84d03e46926e3394b4d2%26id%3Dd13334d318%26e%3Dcc1a53fb05&data=05%7C01%7Csemco%40whoi.edu%7Cd8be5edec9234f67372008db943db7b8%7Cd44c5cc6d18c46cc8abd4fdf5b6e5944%7C0%7C0%7C638266765018760850%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=DKm6OCYWetivLbXSl9iEsWzMwQC%2FRrc0TV28BSq5mFI%3D&reserved=0" originalSrc="https://mbl.us14.list-manage.com/track/click?u=9cefa84d03e46926e3394b4d2&id=d13334d318&e=cc1a53fb05" shash="EARCS7qE+e/CdPbzXEkzPamBFvb0wScVeo/XmfW/ZSGVpjjbMrMInGS8tOVL2CgI83WthK8CcevB4NS6/gwPuspNaoX9imS/4xf7DiIBvZZWLglWZokYkbac8Urlp+nN1kU187YaOe8I43D3Um2iAybuNq1ezC3qEaTjR74eE+k=" target="_blank"><span style="color:windowtext;text-decoration:none"><span style="color:blue"><img border="0" width="24" height="24" style="width:.25in;height:.25in" id="_x0000_i1028" src="https://cdn-images.mailchimp.com/icons/social-block-v2/outline-light-instagram-48.png" alt="Instagram"></span></span></a><o:p></o:p></p>
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<table class="MsoNormalTable" border="0" cellspacing="0" cellpadding="0" width="100%" style="width:100.0%;border-collapse:collapse;-ms-text-size-adjust: 100%;-webkit-text-size-adjust: 100%">
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<td style="padding:3.75pt 7.5pt 3.75pt 6.75pt;-ms-text-size-adjust: 100%;-webkit-text-size-adjust: 100%">
<table class="MsoNormalTable" border="0" cellspacing="0" cellpadding="0" align="left" width="0" style="border-collapse:collapse;-ms-text-size-adjust: 100%;-webkit-text-size-adjust: 100%">
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<td width="24" style="width:.25in;padding:0in 0in 0in 0in;-ms-text-size-adjust: 100%;-webkit-text-size-adjust: 100%">
<p class="MsoNormal" align="center" style="text-align:center"><a href="https://nam02.safelinks.protection.outlook.com/?url=https%3A%2F%2Fmbl.us14.list-manage.com%2Ftrack%2Fclick%3Fu%3D9cefa84d03e46926e3394b4d2%26id%3D75dbe74c28%26e%3Dcc1a53fb05&data=05%7C01%7Csemco%40whoi.edu%7Cd8be5edec9234f67372008db943db7b8%7Cd44c5cc6d18c46cc8abd4fdf5b6e5944%7C0%7C0%7C638266765018760850%7CUnknown%7CTWFpbGZsb3d8eyJWIjoiMC4wLjAwMDAiLCJQIjoiV2luMzIiLCJBTiI6Ik1haWwiLCJXVCI6Mn0%3D%7C3000%7C%7C%7C&sdata=vlpR65dp3lHHNovxaS7e%2FBCnYzoF9yodPLT3%2FzEosK8%3D&reserved=0" originalSrc="https://mbl.us14.list-manage.com/track/click?u=9cefa84d03e46926e3394b4d2&id=75dbe74c28&e=cc1a53fb05" shash="YrTro7D+tK1SL+qXDhEQmXZV7pqqtSnUX3YaFcF4ttAxP6ce/oiBp718k0KMIdr0Gj8sEXcTO+eZNDF8ZNjYxLuKNIkSPiz5wiG+k676zJwGe5sSA3VYYZ0xDSpekFLykurCjXR8ngE/sidC7zD1G3ooQ+RhTgK8CgLAsYkz1fw=" target="_blank"><span style="color:windowtext;text-decoration:none"><span style="color:blue"><img border="0" width="24" height="24" style="width:.25in;height:.25in" id="_x0000_i1027" src="https://cdn-images.mailchimp.com/icons/social-block-v2/outline-light-youtube-48.png" alt="YouTube"></span></span></a><o:p></o:p></p>
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<p class="MsoNormal"><o:p> </o:p></p>
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<table class="MsoNormalTable" border="0" cellspacing="0" cellpadding="0" align="left" width="100%" style="width:100.0%;border-collapse:collapse;-ms-text-size-adjust: 100%;-webkit-text-size-adjust: 100%">
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